Definitely need a constant load path from roof trusses to wall (clips) and wall to the foundation. 48" anchor bolt spacing for the bases is pretty common for what we do here in Indiana and Iowa has the same wind design speeds as us. A lot of times though, based on code uplift, the dead load of the roof will offset the majority of that uplift and not many truss clips or base anchors are needed (hence the seemingly large 4' spacing)
Congrats on building the house OP! Just moved into my newly built house in June. Very excited stuff. I'm surprised you're 6 months in though and still have six months left. The whole process took me 5-6 months total
We just broke ground last week. The first 6 months was the planning and BS to get things set in motion.